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The Journal of the Royal Society of Thailand Quarterly Journal ADVISORY BOARD Pakorn Adulbhan President of the Royal Society Somsak Damronglerd President of the Academy of Science EDITOR-IN-CHIEF Mongkol Dejnakarintra FRI DEPUTY EDITOR Somchai Wongwises FRI EDITORIAL BOARD Krit Chatamara FRI Chayan Picheansoonthon FRI Nibondh Saibejra FRI Piamsak Menasveta FRI Yong Poovorawan FRI Visuth Baimai FRI Somchai Bovornkitti FRI Saichol Ketsa FRI Suda Kiatkamjornwong FRI Suthat Yoksan FRI Sasithon Phukrittayakhamee AFRI Siriwat Wongsiri AFRI Pongsak Siriwong Secretary-General DEPARTMENTAL STAFF Boontham Kranthong Director of Science Division Aree Poldee Literateur, Senior Professional Level Somsong Sakuntanak Literateur, Professional Level Rattikal Sriamphai Literateur, Professional Level Yanut Sai-ngam Literateur, Professional Level Nutthamard Musikacharoen Literateur, Professional Level Anhchana Phaijit-hathai Literateur, Practitioner Level Suguntarat Chamjitchun General Service Officer, Experienced Level MANAGER Supunya Chomchinda Secretary of the Royal Society